Goffinet, B. 1997. Circumscription and phylogenetic trends in the Orthotrichales (Bryopsida). Ph.D. thesis. University of Alberta, Edmonton, Canada.

Goffinet, B. 1989. La végétation épiphytique de la hêtraie du Rurbusch. Bases écologiques nouvelles pour la gestion de la Réserve forestière. Mémoire de licence. Université de Liège, Liège, Belgium.



9. Rozzi, R., F. Massardo, A. Mansilla, F.A. Squeo, E. Barros, T. Contador, M. Frangopulos, E. Poulin, S. Rosenfeld, B. Goffinet, C. González-Weaver, R. MacKenzie, R.D. Crego, F. Viddi, J. Naretto, M.R. Gallardo, J.E. Jiménez, J. Marambio, C. Pérez, J.P. Rodríguez, F. Méndez, O. Barroso, J. Rendoll, E. Schüttler, J. Kennedy, P. Convey, S. Russell, F. Berchez, P.Y.G. Sumida, P. Rundell, A. Rozzi, J. Armesto, M. Kalin-Arroyo & M. Martinic. 2017. Parque Marino Cabo de Hornos — Diego Ramirez. Informe Técnico para la Propuesta de Creación. Programa de Conservación Biocultural Subantártica, Ediciones Universidad de Magallanes, Punta Arenas, Chile. To view the book on-line click on the image.

8. Rozzi, R., L. Lewis, F. Massardo, Y. Medina, K. Moses, M. Ménendez, L. Sancho, P. Vezzani, S. Russell & B. Goffinet. 2012. Ecotourism with a Hand Lens in the Omora Park. 190 p. Photography by Adam M. Wilson and collaborators. Sub-Antarctic Biocultural Conservation Program, Universidad de Magallanes – University of North Texas. Ediciones Universidad de Magallanes.

Book cover7. Goffinet B., R. Rozzi, L. Lewis, W.R. Buck & F. Massardo. 2012. Miniature Forest of Cape Horn/Los Bosques en Miniatura del Cabo de Hornos. Ecotourism with a hand lens/Ecotourismo con lupa. 445 p. University of North Texas Press (Denton, TX) & Ediciones Universidad de Magallanes (Punta Arenas, Chile). Order here   Google scholar

6. Vanderpoorten, A. & B. Goffinet. 2009. Introduction to Bryophyte Biology. 303 p. Cambridge University Press. Cambridge, UK. Order here   Google scholar

5. Goffinet B. & A.J. Shaw (eds.). 2009. Bryophyte Biology (2nd edition). 565 p. Cambridge University Press. Order here   Google scholar

4. Goffinet B., W. R. Buck, P. Massardo & R. Rozzi. 2006. The Miniature Forests of Cape Horn. Fantastico Sur – Universidad de Magallanes. 255 p. Punta Arenas, Chile.

3. Goffinet B., V. Hollowell & R. Magill (eds.). 2004. Molecular Systematic of Bryophytes. Monographs in Systematic Botany from the Missouri Botanical Garden 98: 1-448. Order here  Google scholar

2. Shaw, A.J. & B. Goffinet (eds.). 2000. Bryophyte Biology. 476 pp. Cambridge University Press.   Google scholar

1. Goffinet B. & U. Penz. 1991. Im Wallonischen Venn. German translation of “Promenades en Fagne wallonne” by Frankard et al. (1990). 141 pp. ASBL Hautes-Fagnes, Belgium.


Book chapters and contributions

16. Sánchez-Jardón, L., B. Goffinet & R. Rozzi. 2020. Pioneros Vegetales. Pages 194–195. In: R. Rozzi, F. Morello, F. Massardo & C. Aldunate (eds.), Estrecho de Magallanes: Tres Descubrimientos. Santiago, Chile: Colección Santander, Museo de Chileno de Arte Precolombino. pdf of entire book

15. Rozzi R., M.T. La Valle, S. Russell, B. Goffinet & F. Massardo. 2020. Ecotourism with a hand-lens: a field environmental philosophy experience from the South of the world. Pages: 222–239. In R. Frodeman & E. Brister (eds.), A Guide to Field Philosophy. Case Studies and Practical Strategies (1stedition). Routledge, a Taylor and Francis Group. pdf. Google scholar

14. Goffinet B. & W. R. Buck. 2013. The evolution of body form in bryophytes. Pages 51–90. In: B. Ambrose and M. Purruganan (eds.), The Evolution of Plant Form. Wiley–Blackwell. Annual Plant Reviews 45. pdf   Google scholar

13. Goffinet B., W.R. Buck & A.J. Shaw. 2009. Morphology and classification of the Bryophyta. Pages 55–138. In: B. Goffinet & A.J. Shaw (eds.), Bryophyte Biology, 2nd edition. Cambridge University Press.   Google scholar

12. Goffinet B. 2007. Aphanorrhegma Sull. Physcomitrella Bruch & Schimper, Pyramidula Bridel and Rhachithecium Le Jolis. Pages : 181–182, 194–195, 199 and 468–469, respectively. In: Flora of North America Editorial Committee, eds. 1993+. Flora of North America North of Mexico. New York and Oxford. 27.

11. Vanderpoorten A., B. Goffinet & D. Quandt. 2006. Utility of the internal transcribed spacers of the 18S-5.8S-26S nuclear ribosomal DNA in land plant systematics with special emphasis on Bryophytes. Pages 385–407. In: A.K. Sharma & A. Sharma (eds.), Plant Genome: Biodiversity & Evolution. Volume 2B: Lower plants. Science Publishers, Enfield NH.   Google scholar

10. Goffinet B. 2006. Splachnobryaceae. Pages 159–162. In: P. McCarthy (ed.), Flora of Australia Volume 51: Mosses 1. Australian Biological Resources Study and CSIRO Publishing

9. Goffinet B. 2006. Splachnaceae. Pages 173–181. In: P. McCarthy (ed.), Flora of Australia Volume 51: Mosses 1. Australian Biological Resources Study and CSIRO Publishing.

8. Goffinet B., A.J. Shaw, C.J. Cox, N. J. Wickett, & S. Boles. 2004. Phylogenetic inferences in the Orthotrichoideae (Orthotrichaceae: Bryophyta) based on variation in four loci from all genomes. Pages 270–289. In: B. Goffinet, V. Hollowel & R. Magill (eds.), Monographs in Systematic Botany from the Missouri Botanical Garden 98.

7. Goffinet B., & W.R. Buck. 2004. Systematics of Bryophyta: from molecules to a revised classification. Pages 205–239. In: B. Goffinet, V. Hollowel & R. Magill (eds.), Monographs in Systematic Botany from the Missouri Botanical Garden 98.  Google scholar

6. Sérusiaux E. & B. Goffinet. 2000. Lichens et Bryophytes sur le site de la Makandé. Pages 68–89. In: F. Hallé (ed.), Biologie d’une canopé de forêt équatoriale – IV. Rapport de la mission du radeau des cimes à la Makandé, forêt des Abeilles, Gabon. Janvier – Mars 1999. Pro Natura International.

5. Goffinet B. 2000. Origin and phylogenetic relationships of the Bryophyta. Pages 124–149. In: A. J. Shaw & B. Goffinet (eds.), Bryophyte Biology. Cambridge University Press.  Google scholar

4. Buck W.R. & B. Goffinet. 2000. Morphology and classification of mosses. Pages 71–123. In: A. J. Shaw & B. Goffinet (eds.), Bryophyte Biology. Cambridge University Press.  Google scholar

3. Goffinet B., and T. Goward. 1998. Is Nephroma silvae-veteris the cyanomorph of Lobaria oregana? Insights from molecular, chemical and morphological characters. Pages 41–52. In: M.G., Glenn, R.C. Harris & M.S. Cole (eds.), Lichenographia Thomsoniana: North American Lichenology in Honor of John W. Thomson. Mycotaxon Ltd, Ithaca NY.

2. Goffinet, B. & D.H. Vitt. 1998. A revised classification of the Orthotrichales (Bryopsida) based on comparative morphology and a rbcL based phylogeny. Pages 143–159. In: J.W. Bates, N.W. Ashton & J.G. Duckett (eds.), Bryology for the Twenty-First Century. Proceedings of the Centenary Symposium of the British Bryological Society. Leeds: Maney and The British Bryological Society.  Google scholar

1. Vitt, D.H., B. Goffinet, & T.A. Hedderson. 1998. The ordinal classification of the mosses: Questions and answers for the 1990’s. Pages 113–123. In: J.W. Bates, N.W. Ashton & J.G. Duckett (eds.), Bryology for the Twenty-First Century. Proceedings of the Centenary Symposium of the British Bryological Society. Leeds: Maney and The British Bryological Society. Google scholar


Refereed Publications

172. Dong S., J. Yu, L. Zhang, B. Goffinet & Y. Liu. Phylotranscriptomics of liverworts: revisiting the backbone phylogeny and ancestral gene duplications. Annals of Botany accepted.

171. Rozzi R., C. S. Quilodrán, E. Botero-Delgadillo, R.D. Crego, C. Napolitano, O. Barroso, R. MacKenzie, J.C. Torres-Mura, C. Bravo, S. Ippi, V. Quirici, J. Rivero-de-Aguilar, C.G. Suazo, B. Goffinet, B. Kempenaers, E. Poulin & R.A. Vásquez. 2022. Aphrastura subantarctica: the Subantarctic Rayadito, a new bird species on the Southernmost islands of the Americas. Scientific Reports 12: 13957. pdf

170. Liu Y., S. Wang, L. Li, T. Yang, S. Dong, T. Wei, S. Wu, Y. Liu, Y. Gong, X. Feng, J. Ma, G. Chang, J. Huang, Y. Yang, H. Wang, M. Liu, Y. Xu, H. Liang, J. Yu, Y. Cai, Z. Zhang, Y. Fan, W. Mu, S. Kumar Sahu, S. Liu, X. Lang, L. Yang, N. Li, S. Habib, Y. Yang, A.J. Lindstrom, P. Liang, B. Goffinet, S. Zaman, Jill. L. Wegrzyn, D. Li, J. Liu, J. Cui, E.C. Sonnenschein, X. Wang, J. Ruan, J.-Y. Xue, Z.-Q. Shao, C. Song, G. Fan, Z. Li, L. Zhang, J. Liu, Z.-J. Liu, Y. Jiao, X.-Q. Wang, H. Wu, E. Wang, M. Lisby, H. Yang, J. Wang, X. Liu, X. Xu, N. Li, P.S. Soltis, Y. Van de Peer, D.E. Soltis, X. Gong, H. Liu, S. Zhang. 2022. The cycad genome and early evolution of seed plants. Nature Plants 8: 389–401. pdf  Google Scholar

169. Patiño J., I. Bisang, B. Goffinet, L. Hedenäs, S. McDaniel, S. Pressler, M. Stech, C. Ah-Peng, A. Bergamini, R.T. Caners, C. Cargill, N. Cronberg, J. Duckett, S. Eppley, N. Fenton, K. Fisher, J. M. González-Mancebo, M. Hasebe, J. Heinrichs†, K. Hylander, M.S. Ignatov, J. Martínez-Abaigar, N. Medina, R. Medina, D. Quandt, S. Rensing, K. Renzaglia, M. Renner, R. M. Ros, A. Schäfer-Verwimp, J. Carlos Villarreal & A. Vanderpoorten. 2022. Unveiling the nature of a miniature world: A horizon scan of fundamental questions in bryology. Journal of Bryology 41: 1–34. pdf  Google Scholar

168. Yu J., Y. Cai, Y. Zhu, Y. Zeng, S. Dong, K. Zhang, S. Wang, L. Li, B. Goffinet, H. Liu & Y. Liu. 2022. Chromosome-level genome assemblies of two Hypnales mosses reveal high intergeneric synteny. Genome Biology and Evolution 14: evac020. pdf  Google scholar

167. Dong S., H.-L. Li, B. Goffinet & Y. Liu. 2022. Exploring the impact of RNA editing on mitochondrial phylogenetic analyses in liverworts, an early land plant lineage. Journal of Systematics and Evolution 60: 16–22. pdf  Google scholar

166. Simon A., B. Goffinet, L.S. Wang, T. Spribille, T. Goward, T. Pystina, N. Semenova, N.V. Stepanov, B. Moncada, R. Lücking, N. Magain & E. Sérusiaux. 2022. Global phylogeny and taxonomic reassessment of the genus Dendriscosticta (Ascomycota: Peltigerales). Taxon 71: 256–287 pdf  Google scholar

165. Widhelm T.J., F. Grewe, B. Goffinet, M. Wedin, T. Goward, L.F. Coca, I. Distefano, A. Košuthová & H.T. Lumbsch. 2021. Phylogenomic reconstruction addressing the Peltigeralean backbone (Lecanoromycetes, Ascomycota). Fungal Diversity 110: 59–73pdf  Google scholar

164. Collart F., J. Wang, J. Patino, A. Hagborg, L. Söderström, B. Goffinet, N. Magain, O.J. Hardy & A. Vanderpoorten. 2021. Macroclimatic structuring of spatial phylogenetic turnover in liverworts. Ecography 43: 1474–1485. pdf  Google scholar

163. Frangedakis E., F. Guzman-Chavez, M. Rebmann, K. Markel, Y. Yu, A. Perraki, S. Wai Tse, Y. Liu, J. Rever, S. Sauret-Gueto, B. Goffinet, H. Schneider & J. Haseloff. 2021. A system for rapid genome engineering and hyper-expression in plastids. ACS Synthetic Biology 10: 1651–1666. pdf Google scholar

162. Dong S., S. Zhang, L. Zhang, H. Wu, B. Goffinet & Y. Liu. 2021. Plastid genomes and phylogenomics of the early land plant liverworts (Marchantiophyta): conserved genome structure and highest plastid substitution rates among three genetic compartments. Molecular Phylogenetics and Evolution 161: 107171. pdf   Google scholar

161. Patel N., R. Medina, M. Johnson & B. Goffinet. 2021. Karyotypic diversity and cryptic speciation: Have we vastly underestimated moss species diversity? Bryophyte Diversity and Evolution 43: 150–163. pdf  Google scholar

160. Frye H.A., Z. Muscavitch & B. Goffinet. 2021. Discovery of epiphytic lichens in Connecticut suggests novel introduction and reintroduction via horticultural practices. The Bryologist  124: 191–197. pdf  Google scholar

159. Rahmatpour N., N.V. Perera, V. Singh, J.L. Wegrzyn & B. Goffinet. 2021. High gene space divergence contrasts with frozen vegetative architecture in the moss family Funariaceae. Molecular Phylogenetics and Evolution 154: 106965. pdf  Google scholar

158. Yu J., L. Li, S. Wang, S. Dong, Z. Chen, N. Patel, B. Goffinet, H. Liu & Y. Liu. 2020. Draft genome of the aquatic moss Fontinalis antipyretica (Fontinalaceae, Bryophyta). Gigabyte 1 pdf

157. Sánchez-Jordón L., B. Goffinet & R. Rozzi. 2020. Los colonizadores vegetales del estrecho de Magallanes y su función indicadora en el cambio climático. Adumbrationes as Summae Editionem 94: 1–14. (with photographs by J.F. Soza & B. Goffinet).

156. Sun G., S. Bai, Y. Guan, Q. Wang, S. Wang, Y. Liu, H. Liu, B. Goffinet, Y. Zhou, M. Paoletti, X. Hu, F. Haas, N. Fernendez-Pozo, A. Czyrt, H. Sun, S. Rensing & J. Huang. 2020. Are fungi-derived genomic regions related to antagonism toward fungi in mosses? New Phytologist 228: 1169–1175. pdf  Google scholar

155. Lindgren H., B. Moncada, R. Lücking, N. Magain, A. Simon, B. Goffinet, E. Sérusiaux, M.P. Nelsen, J. Mercado-Díaz, T. Widhelm & T. Lumbsch. 2020. Cophylogenetic patterns in algal symbionts correlate with repeated symbiont switches during diversification and geographic expansion of lichen-forming fungi in the genus Sticta (Ascomycota, Peltigeraceae). Molecular Phylogenetics and Evolution 150: 106860. pdf  Google scholar

    154. Mackenzie R., O. Vidal, S. Rosenfeld, T. Contador, O. Barroso, B. Goffinet, F. Massardo, P. Arce-Johnson & R. Rozzi. 2020. Flora vascular y ausencia de especies exóticas en el archipiélago Diego Ramírez (56°31’S), Chile. Anales del Instituto de la Patagonia 48(3): 138–148. pdf  Google scholar

    153. Goffinet B., J.J. Engel, M. von Konrat, R. MacKenzie, T. Contador, S. Rosenfeld, O. Barroso & R. Rozzi. 2020. Bryophyte records from Isla Gonzalo, Diego Ramirez Islands, Americas’ deep southern ocean archipelago. Anales del Instituto de la Patagonia 48(3): 127–138. pdf  Google scholar

    152. Marambio J., S. Rosenfeld, J. P. Rodríguez, F. Méndez, T. Contador, R. Mackenzie, B. Goffinet, R. Rozzi & A. Mansilla. 2020. Siete nuevos registros de macroalgas para el archipiélago Diego Ramírez (56°31’S): el valor del nuevo parque marino como sumidero de carbono y conservación de la biodiversidad subantártica. Anales del Instituto de la Patagonia 48(3): 99–111. pdf  Google scholar

    151. Contador T., J. Rendoll, R. Mackenzie, S. Rosenfeld, O. Barroso, R. Rozzi, B. Goffinet, J. Kennedy & P. Convey, P. 2020. Comunidades de Invertebrados Terrestres en el Archipiélago Diego Ramírez (56°, 31’s) y sus Afinidades con las Islas Sub-Antárticas del Océano Austral. Anales del Instituto de la Patagonia 48(3): 83–98. pdf  Google scholar

    150. Rozzi R., R.D. Credo, T. Contador, E. Schüttler, S. Rosenfeld, R. MacKenzie, O. Barroso, E.A. Silva-Rodríguez, X. Álvarez Bustos, A. Silva, I. Ramírez, J. Mella, J. Herreros, J. Rendoll-Cárcamo, J. Marambio, J. Ojeda, F. Méndez, K.P. Moses, J. Kennedy, S. Russell, B. Goffinet, F. Aguirre, L. Sánchez-Jardón, E. Barros, R.A. Vásquez, E. Poulin, F. Squeo, J.J. Armesto, A. Mansilla & F. Massardo. 2020. Extensión de la Red de Estudios Socio-Ecológicos a Largo Plazo (LTSER-Chile) en la Reserva de la Biosfera Cabo de Hornos y el Nuevo Parque Marino Islas Diego Ramírez-Paso Drake (Extension of the long term socio-ecological research network (LTSER-Chile) in the Cape Horn Biosphere Reserve and the new marine park Diego Ramirez Islands Drake passage). Anales del Instituto de la Patagonia 48(3): 45–81. pdf  Google scholar

    149. Kirbis A., M. Waller, M. Ricca, Z. Bont, A. Neubauer, B. Goffinet & P. Szövényi. The transcriptomic landscape of differential sporophyte development in two mosses, Physcomitrium (Physcomitrellapatens and Funaria hygrometrica. Frontiers in Plant Science 11: 747. pdf  Google scholar

    148. Simon A., R. Lücking, B. Moncada, J.A. Mercado-Díaz, F. Bungartz, M. Cáceres, E. Gumboski, S. Maria de Azevedo Martinsi, D. Parker & B. Goffinet. 2020. Emmanuelia, a new genus of lobarioid lichen-forming fungi (lichenized Ascomycota: Peltigerales). Plant and Fungal Systematics 65: 76–94. pdf  Google scholar

    147. Magain N., B. Goffinet, A. Simon, J. Seelan Sathiya, I. Medeiros, F. Lutzoni & J. Miadlikowska. 2020. Peltigera serusiauxii, a new species in section Polydactylon from Papua New Guinea and Malaysia (Lecanoromycetes, Ascomycota). Plant and Fungal Systematics 65: 139–146. pdf Google scholar

    146. Robertson M.W., N.J. Russo, S.J. McInnes, B. Goffinet & J.E. Jiménez. 2020. Potential dispersal of tardigrades by birds through endozoochory: evidence from sub-Antarctic White-bellied Seedsnipe (Attagis malouinus). Polar Biology 43: 899–902. pdf. Google scholar

    145. Alonso Garcia M., Villarreal J.C., K. McFarland & B. Goffinet. 2020. Population genomics confirms extreme sex ratio of a clonal bryophyte. Frontiers in Plant Science 11: 495. pdf  Google scholar

    144. Rensing S.A., B. Goffinet, R. Meyberg, S.-Z. Wu & M. Bezanilla. 2020. The moss Physcomitrium (Physcomitrella) patens: a model organism for non-seed plants. The Plant Cell 32: 1361–1376. pdf Google scholar

    143. Russo N.J., M. Robertson, R. MacKenzie, B. Goffinet & J.E. Jimenez. 2020. Evidence of targeted consumption of mosses by birds in Sub-Antarctic South America. Austral Ecology 45: 399–403.  pdf Google scholar

    142. Zhang J., X.-X. Fu, R.-Q. Li, X. Zhao, Y. Liu, M.-H. Li, A. Zwaenepoel, H. Ma, B. Goffinet, Y.-L. Guan, J.-Y. Xue, Y.-Y. Liao, Q.-F. Wang, Q.-H. Wang,Q.-H. Wang, J.-Y. Wang, G.-Q. Zhang, Z.-W. Wang, Y. Jia, M.-Z. Wang, S.-S. Dong, J.-F. Yang, Y.-N. Jiao, Y.-L. Guo, H.-Z. Kong, A.-M. Lu, S.-Z. Zhang, Y. Van de Peer, Z.-J. Liu & Z.-D. Chen. 2020. The hornwort genome and early land plant evolution. Nature Plants 6: 107118. pdf Google scholar

    141. Dong S., C. Zhao, S. Zhang, L. Zhang, H. Wu, R. Zhu, Y. Jia, B. Goffinet & Y. Liu. 2019. Mitochondrial genomes of the early land plant lineage liverworts (Marchantiophyta): conserved genome structure, and ongoing low frequency recombination. BMC Genomics 20: 953. pdf  Google scholar

    140. Dong S., C. Zhao, S. Zhang, H. Wu, W. Mu, T. Wei, N. Li, H. Liu, J. Cui, R. Zhu, B. Goffinet & Y. Liu. 2019. The amount of RNA editing sites in liverwort organellar genes is correlated with GC content and nuclear PPR protein diversity. Genome Biology and Evolution 11: 3233–3239. pdf Google scholar

    139. Medina R., M. G. Johnson, Y. Liu, N.J. Wickett, A.J. Shaw & B. Goffinet. 2019. Phylogenomic delineation of Physcomitrium (Bryophyta: Funariaceae) based on nuclear targeted exons and their flanking regions rejects the retention of Physcomitrella, Physcomitridium and Aphanorrhegma. Journal of Systematics and Evolution 57: 404–417. pdf. Google scholar

    138. Widhelm T.J., F. Grewe, J.P. Huang, J. Mercado, B. Goffinet, R. Lücking, I. Schmitt, B. Moncada, R. Mason-Gamer & H.T. Lumbsch. 2019. Multiple historical processes obscure phylogenetic relationships in a taxonomically difficult group (Lobariaceae, Ascomycota). Scientific Reports 9: 8968. pdf Google scholar

    137. Liu Y., M.G. Johnson, C.J. Cox, R. Medina, N. Devos, A. Vanderpoorten, L. Hedenäs, N.E. Bell, J. R. Shevock, B. Aguero, D. Quandt, N. J. Wickett, A. J. Shaw & B. Goffinet. 2019. Resolution of the backbone phylogeny of mosses using targeted exons from organellar and nuclear genomes. Nature Communications 10: 1485. pdf Google scholar

    136. Dong S., Q. He, S. Zhang, H. Wu, B. Goffinet & Y. Liu. 2019. The mitochondrial genomes of Bazzania tridens and Riccardia planiflora further confirms conservative evolution of mitogenomes in liverworts. The Bryologist 122: 130–139. pdf  Google scholar

    135. Medina R., F. Lara, B. Goffinet, R. Garilleti & V. Mazimpaka. 2019. On the priority of Orthotrichum cylindrocarpum Lesq. over O. coulteri Mitt. and Lesquereux’s early vindication of an autonomous American bryology. Taxon 68: 137–141. pdf  Google scholar

    134. Magain N., C. Truong, T. Goward, D. Niu, B. Goffinet, E. Sérusiaux, O. Vitikainen, F. Lutzoni & J. Miadlikowska. 2018.. Species delimitation at a global scale reveals high species richness with complex biogeography and patterns of symbiont association in Peltigera section Peltigera (lichenized Ascomycota: Lecanoromycetes). Taxon 67: 836–870. pdf  Google scholar

    133. Liu D., B. Goffinet, X. Wang, J.-S. Hur, M. Yang, H. Shi, Y. Zhang, C. An & L. Wang. 2018. Another lineage of basidiolichen in China, with one new species and new record in the genera Dictyonema and Lichenomphalia (Agaricales: Hygrophoraceae). Mycosystema 37: 849–864. pdf  Google scholar

    132. Lewis L.R., C. Gottschalk Druschke, C. Saldías, R. Mackenzie, B. Goffinet & R. Rozzi. 2018. Cultivando un jardín de nombres en los bosques en miniature del Cabo de Hornos: extensión de la conservación biocultural y la ética a seres vivos poco percibidos (Cultivating a Garden of Names in the Miniature Forests of Cape Horn: an extension of biocultural conservation and ethics to under-perceived beings). Magallania 46: 103–123. pdf Google scholar

    131. Laenen B, J. Patiño, A. Hagborg, A. Désamoré, J. Wang, A. J. Shaw, B. Goffinet & A. Vanderpoorten. 2018. Evolutionary origin of the latitudinal diversity gradient in liverworts. Molecular Phylogenetics and Evolution 127: 606–612. pdf  Google scholar

    130. Simon A., J. Di Meglio, T. Goward, K. Dillman, T. Spribille & B. Goffinet. 2018. Sticta torii sp. nov., a remarkable lichen of high conservation priority from northwestern North America. Graphis Scripta 30: 105–114. pdf. Google scholar

    129. Widhelm T.J., F.R. Bertoletti, M.J. Asztalos, J.A. Mercado-Díaz, J.-P. Huang, B. Moncada, R. Lücking, N. Magain, E. Sérusiaux, B. Goffinet, N. Crouch, R. Mason-Gamer & H. T. Lumbsch. 2018. Oligocene origin and drivers of diversification in the genus Sticta (Lobariaceae, Ascomycota). Molecular Phylogenetics and Evolution 126: 58–73. pdf  Google scholar

    128. Liu D., B. Goffinet, D. Ertz, A. De Kesel, Z.G. Qian, X.Y. Wang, H.X. Shi, Y.Y. Zhang, J.W. Li, X. Ye, J.S. Hur & L.S. Wang. 2017 (2018). Circumscription and phylogeny of the Lepidostromatales (lichenized Basidiomycota) following discovery of new species from China and Africa. Mycologia 109: 730–748. pdf. Google scholar

    127. Simon A., B. Goffinet, N. Magain & E. Sérusiaux. 2018. High diversity, high insular endemism and recent origin in the lichen genus Sticta (lichenized Ascomycota, Peltigerales) in Madagascar and the Mascarene archipelago. Molecular Phylogenetics and Evolution 122: 15–28. pdf. Google scholar

    126. Dong S., J.-Y. Xue, S. Zhang, L. Zhang, H. Wu, Z. Chen, B. Goffinet & Y. Liu. 2018. Complete mitochondrial genome sequence of Anthoceros angustus: conservative evolution of mitogenome in hornworts. The Bryologist 121: 14–22. pdf. Google scholar

    125. Medina R., M. Johnson, Y. Liu, N. Wilding, T. Hedderson, N. Wickett & B. Goffinet. 2018. Evolutionary dynamism in bryophytes: Phylogenomic inferences confirm rapid radiation in the family Funariaceae. Molecular Phylogenetics and Evolution 120: 240–247. pdf. Google scholar

    124. Lücking, B. Moncada, B. McCune, E. Farkas, B. Goffinet, D. Parker, J. L. Chaves, L. Lőkös, P. R. Nelson, T. Spribille, S. Stenroos, T. Wheeler, A. Yanez-Ayabaca, K. Dillman, O. T. Gockman, T. Goward, J. Hollinger, E. A. Tripp, J. Villella, W. R. Álvaro-Alba, C. Julio Arango, M. E. S. Cáceres, L. Fernando Coca, C. Printzen, C. Rodríguez, K. Scharnagl, R. Rozzi, E. Soto-Medina & L.  S. Yakovchenko. 2017. Pseudocyphellaria crocata (Ascomycota: Lobariaceae) in the Americas reveals to be ten species, and none of them is Pseudocyphellaria crocata. The Bryologist 120: 441–500. pdf. Google scholar

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    32. Fife A. & B. Goffinet. 2003. The identity of Tayloria maidenii Broth. The Bryologist 106: 309-310.  pdf  Google Scholar

    31. Goffinet B. 2002 (2003). Orthotrichum sprucei Mont., a European endemic discovered in Kazakhstan. Arctoa 11: 27–30.  Google Scholar

    30. Goffinet B. & R. Shannon. 2002. On the occurrence of an extensive but deciduous prostome in Orthotrichum obtusifolium. The Bryologist 105: 349–352.  pdf  Google Scholar

    29. Goffinet B. & A.J. Shaw. 2002. Independent origins of cleistocarpy in the Splachnaceae: analyses of cpDNA sequences and polyphyly of the Voitioideae (Bryophyta). Systematic Botany 27: 203–208.  pdf  Google Scholar

    28. Vanderpoorten A., A.J. Shaw & B. Goffinet. 2001. Testing controversial alignments in Amblystegium and related genera (Amblystegiaceae: Bryopsida). Evidence from rDNA ITS sequences. Systematic Botany 26: 470–479.  pdf  Google Scholar

    27. Goffinet B, C.J. Cox, A.J. Shaw & T.A. Hedderson. 2001. The Bryophyta (Mosses): Systematic and evolutionary inferences from a rps4 gene (cpDNA) phylogeny. Annals of Botany 87: 191–208.  pdf  Google Scholar

    26. Buck W.R., B. Goffinet & A.J. Shaw. 2000. Novel relationships in pleurocarpous mosses as revealed by cpDNA sequences. The Bryologist 103: 774-789.  pdf  Google Scholar

    25. Goward T., & B. Goffinet. 2000. Peltigera chionophila, a new lichen (Ascomycetes) from the western cordillera of North America. The Bryologist 103: 493–498.  pdf  Google Scholar

    24. Shaw A.J. & B. Goffinet. 2000. Molecular evidence of reticulate evolution in the peatmosses (Sphagnum L.), including S. ehyalinum n. sp. The Bryologist 103: 357–374.  pdf  Google Scholar

    23. Cox, C., B. Goffinet, A.E. Newton, A.J. Shaw & T.A.J. Hedderson. 2000. Phylogenetic relationships among the diplolepideous-alternate mosses (Bryidae) inferred from nuclear and chloroplast DNA sequences. The Bryologist 103: 224–241.  pdf  Google Scholar

    22. Goffinet B. & C. Cox. 2000. Phylogenetic relationships among basal-most arthrodontous mosses with special emphasis on the evolutionary significance of the Funariineae. The Bryologist 103: 212–223.  Google Scholar

    21. Newton A.E., C. Cox, J. G. Duckett , J. Wheeler, B. Goffinet, T.A.J. Hedderson & B.D. Mishler. 2000. Evolution of the major moss lineages. The Bryologist 103: 187–211.  Google Scholar

    20. Buck W.R., B. Goffinet & A.J. Shaw. 2000. Testing morphological concepts of orders of pleurocarpous mosses (Bryophyta) using phylogenetic reconstructions based on trnL-trnF and rps4 sequences. Molecular Phylogenetics and Evolution 16: 180–198.  Google Scholar

    19. Goffinet B. & H. Greven. 2000. Grimmia indica (Grimmiaceae), a new combination. Journal of Bryology 22: 141.  Google Scholar

    18. Goffinet B., A.J. Shaw, L.E. Anderson & B.D. Mishler. 1999. Peristome development in mosses in relation to systematics and evolution. V. Orthotrichaceae. The Bryologist 102: 581–599.  Google Scholar

    17. Goffinet B. & J. Miadlikowska. 1999. Peltigera phyllidiosa (Peltigeraceae, Lichenized Ascomycota), a new morphological species from Eastern North America corroborated by ITS sequences. The Lichenologist 31: 247–256. pdf   Google Scholar

    16. Goffinet B. 1999. Brachymitrion immersum (Splachnaceae, Musci) a new species from Cameroon. The Bryologist 102: 108-111.  pdf  Google Scholar

    15. Goffinet B., R.J. Bayer & D.H. Vitt. 1998. Circumscription and phylogeny of the Orthotrichales (Bryopsida) inferred from rbcL sequence analysis. American Journal of Botany: 85: 1324–1337.  pdf  Google Scholar

    14. Hodgetts N. & B. Goffinet. 1998. Hypnodontopsis mexicana (Thér.) H. Rob. (Rhachitheciaceae), a genus and species new to Africa. Journal of Bryology 20: 251–252.  Google Scholar

    13. Goffinet B. 1998. Jonesiobryum sphaerocarpum Allen and Pursell (Musci: Rhachitheciaceae) new to the Central African Republic. Journal of Bryology 20: 242–243.  Google Scholar

    12. Goffinet B. 1998. A reconsideration of the affinities of Kleioweisiopsis, Pleurozygodontopsis, Trigonodictyon , and the Microtheciellaceae (Bryopsida, Orthotrichales). Journal of Bryology 20: 69–81.  Google Scholar

    11. Goffinet B. 1997 (1998). The Rhachitheciaceae: Revised generic circumscription and ordinal affinities. The Bryologist 100: 425–439.  pdf  Google Scholar

    10. Goffinet B. & R.J. Bayer. 1997. Characterization of mycobionts of photomorph pairs in the Peltigerineae (lichenized Ascomycetes) based on ITS sequences of specifically amplified fungal ribosomal DNA. Fungal Genetics and Biology 21: 228–237. pdf   Google Scholar

    9. Goffinet B. 1996. Cardotiella elimbata (Thér.) comb. nov., and C. appendiculata new to Mauritius (Musci, Orthotrichaceae). The Bryologist 99: 390–396.  pdf  Google Scholar

    8. Goward T., B. Goffinet & O. Vitikainen. 1995. Synopsis of the genus Peltigera (Lichenes, Ascomycotina) in British Columbia with a key to the North American species. Canadian Journal of Botany 73 (1): 91–111. pdf   Google Scholar

    7. Goffinet B. & R. Hastings. 1995. Two new sorediate taxa of Peltigera (Lichenized Ascomycetes). The Lichenologist 27: 43-58.   Google Scholar

    6. Goffinet B. & R. Hastings. 1994. The lichen genus Peltigera (L.) Willd. in Alberta. Provincial Museum of Alberta Natural History Occasional Paper No. 21: i-vi + 1–54.  Google Scholar

    5. Goffinet B., E. Sérusiaux & P. Diederich. 1994 (1995). Le genre Peltigera (Lichenes) en Belgique et au Grand-Duché de Luxembourg. Belgian Journal of Botany 127: 184–206. pdf  Google Scholar

    4. Goffinet B. 1994. Floristic notes on the lichen and lichenicolous fungi flora of Alberta, Canada. Mycotaxon 51: 1–4.  Google Scholar

    3. Goffinet B. 1993. Taxonomic and floristic notes on neotropical Macromitrioideae (Orthotrichaceae). Tropical Bryology 7: 149–154.  Google Scholar

    2. Goward T. & B. Goffinet. 1993. Nephroma silvae-veteris, a new lichen (Ascomycotina) from the Pacific Northwest of North America. The Bryologist 96: 242–244.  Google Scholar

    1. Goffinet B. 1992. Deux lichens nouveaux pour la Belgique: Bryoria subcana (Nyl. ex Stizenb.) Brodo & D. Hawksw. et Micarea melaena (Nyl.) Hedl. Dumortiera 51: 19–21.


    Distribution records and other non-reviewed publications

    Distribution records and other non-reviewed publications

    12. Villarreal J.C., B. Goffinet, M. von Konrat, M. Favreau, N. Fenton & S. Schuette. 2021. Program and abstracts of “BL2021 Bryophytes, lichens, and northern ecosystems in a changing world” of the joint virtual meeting of the International Association of Bryologists, American bryological and Lichenological Society, Canadian Botanical Association and Société québequoise de bryologie. Université Laval, Québec, Canada. Special issue of the Bryological Times 152: 1–178.

    11. Cole T.C.H., H.H. Hilger, J.B. Bachelier, B. Goffinet, P.F. Stevens, N.M. Shiyan, S.L. Zhygalova & S.L. Mosyakin. Spanning the Globe – The Plant Phylogeny Poster (PPP) Project. Ukrainian Botanical Journal: Short Communication 78(3): 235–241.

    q0. Kapoor M., J.G. Duckett, S. Rensing & B.Goffinet. 2021. Editorial: Highlights 1 of IAB IMOSS SEB 2019 Joint Conference. Frontiers in plant Sciences 12: 694765.

    9. Magain N., D. Ertz, B. Goffinet & P. Diederich. 2020. Forty-five years of lichenology: a tribute to Emmanuël Sérusiaux. Plant and Fungal Systematics 65: 2–12.

    8. von Konrat M., B. Goffinet, Y. Rodriquez, B. McDonald, C. Zapata, M. Enkhbayer, E. Arnett, J. Harris & J. Martinec. 2019. Abstracts from the 2019 joint conference between the International Association of Bryologists, the Spanish Bryological Society, and the International Molecular Moss Science Society. Special issue of the Bryological Times 49: 1–100.

    7. Rozzi R., L. Lewis, F. Massardo & B. Goffinet. 2015. Luupidega ökoturism samblafloora väärtustamiseks: ülekutse Lõunast. [Ecotourism with a hand-lens to appreciate the bryoflora: an invitation from the South.] Samblasõber 18: 2–13. pdf

    6. Goffinet B. 2004. Bibliography of systematic and population genetic studies of Bryophytes based on DNA data. III. Cryptogamie, Bryologie 25: 385–390.  Google Scholar

    5. Goffinet B. 2003. Bibliography of systematic and population genetic studies of Bryophytes based on DNA data. II. Cryptogamie, Bryologie 24: 181–185.  Google Scholar

    4. Goffinet B., R. Rosentreter & E. Sérusiaux. 2001. A second locality for Xanthoparmelia idahoensis Hale, an endangered vagrant lichen, new to Canada. Evansia 18: 58–59.  Google Scholar

    3. Goffinet B. & N.P. Hax. 2001. Bibliography of “molecular systematic” studies of bryophytes. I. 1985-2000. Cryptogamie — Bryologie 22: 1–7.  Google Scholar

    2. Goffinet B. & T.A.J. Hedderson. 2000. “Evolutionary biology of the Bryopsida: A synthesis”. Introduction. The Bryologist 103: 185–186.

    1. Goffinet B. 1992. The North American distribution of Peltigera retifoveata. Evansia 9: 49–51.


    Electronic dissemination

    2. Cole T.C.H., H.H. Hilger & B. Goffinet. 2019. Bryophyte Phylogeny Poster (BPP) PeerJ Preprints 7:e27571v3 (translated in 20 other languages and available on

    1. Goffinet B., W.R. Buck & A.J. Shaw. Constantly updated. Classification of the Bryophyta.


    Book reviews

    11. Medina R. & B. Goffinet. 2014. Better shoes alone don’t get you to your destination.  Review of Zander, R. H. 2013. A Framework for Post-Phylogenetic Systematics. The Bryologist 117: 431–433.

    10. Goffinet B. 2003. Review of Biologie der Moose. By Frahm J.-P. The Bryologist 106: 494-495.  Google Scholar

    9. Goffinet B. 2002. Review of Lexikon deutschsprachiger Bryologen. By Frahm J.-P. and J. Eggers. The Bryologist 105: 297-298

    8. Goffinet B. 1999. Review of American Arctic Lichen. 2. The Microlichens. By J.W. Thomson. The Bryologist 102: 345-346.

    7. Goffinet B. 1998. Review of Macrolichens of the Pacific Northwest. By B. McCune and P. Geiser. The Bryologist 101: 478.

    6. Goffinet B. 1998. Review of Flora of the Guianas. Series C. Bryophytes. By Görts-van Rijn A.R.A. (ed.). Nordic Journal of Botany 18: 322.

    5. Goffinet B. 1997. Review of Niebla and Vermilacinia (Ramalinaceae) in California and Baja California. By R.W. Spjut (1997). Plant Science Bulletin 43: 83-85.

    4. Goffinet B. 1996. Review of Die Moos- und Farnpflanzen Europas. By Frey, W., J.-P. Frahm, E. Fischer & W. Lobin (1995). The Bryologist 99: 379.

    3. Goffinet B. 1994. Review of The Lichens of Italy. An Annotated Catalogue. By P.L. Nimis (1993). Vegetatio 116: 173.

    2. Goffinet B. 1994. Review of Monographic Studies in Orthotrichum (Musci). By J. Lewinsky (1993). The Bryologist 97: 464-465.

    1. Goffinet B. 1993. Review of Populationsbiologie der Pflanzen – Grundlagen – Probleme – Perspektiven. By K.M Urbanska (1992). Plant Science Bulletin 39: 18-20.



    1. Right Below Your Feet. A beautiful evolutionary success story. An exhibit of nine 6-foot metal prints of mosses from North and South America, planned and produced by the Connecticut State Museum of Natural History in collaboration B. Goffinet and Mark Smith (Macroscopic Solutions). University of Connecticut, Fall 2020-Spring 2021.